Tongue and groove flooring has been around since the early 1800's some houses of post industrial age t&g wood was done by hand using mortison and tendon type interlocking. Since t&g flooring was either fastend down using a hand drive or a cleat style fastener The reason for the 1.5 inch wide by 3/8,5/8, or commercial 3/4" thicknesses because it was easier to get a tight fit and wood was not always the straightest making it easier to get the bows out when nailing down, The t&g are milled in such a way as the wood needs to forced together to ensure a tight fit. Hence the reason why today we us an air stapler that needs to be hit with a mallet to ensure a tight fit.
If you have a board that has been faced nailed that is not around the parameter of the room, it is most likely a board that has become loose over the years because older houses in climates that have seasonal weather expand and contract causing the nails to lossen up over time. The other could be that a board was replaced because of damage for some reason or another, when the board was replaced the new board had the tongue removed and the bottom wood strip removed on the groove side so the board could be installed and the face would be nailed down.
Usually after the hardwood floors were fixed of squeaking and damage caused by rodents, water stains, or heavy objects the floors wouks be sanded down to bare wood nail holes wood be filled as well as cracks or knots that have dried out. Then you would sand the floor atleast 2 more times depending on the hardness of the wood. A maple, walnut, Mahogany, Brazilian cherry (jatoba wood) some of your harder woods that may need to be sanded in steps using finer grits to take out the scratches ( swirls caused by the sander) Oak is one of the woods that usually only needs to be sanded a few times to get the floor ready so it can be sealed with some sort of protective coating.
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